一、 简介

杨贤鹏，男，汉族，理学博士，安徽宿松人，1989年11月生。

工作经历:

2023/12-至今 山东师范大学生命科学学院  副教授，硕士生导师

2019/07-2023/12 山东师范大学生命科学学院 讲师，硕士生导师

2017/07-2019/06 中国科学院武汉植物园 博士后

教育经历:

2013/09-2017/06 中国科学院武汉植物园 植物学 博士

2010/09-2013/06 中国科学院近代物理研究所 生物物理学 硕士

2006/09-2010/06 吉林大学 生物技术 学士

二、 研究方向

长期从事植物表皮脂质（蜡质、角质和木栓质）的合成调控分子机制研究，并探讨表皮脂质在植物应答环境非生物胁迫中的功能，涉及植物材料包括小麦、藜麦、拟南芥和盐芥等。

三、 承担项目

1. 主持国家自然科学基金青年基金项目：乙酰辅酶A羧化酶ACC1参与拟南芥蜡质合成响应盐胁迫的功能解析（32100207），2022-2024，30万，已结题；

2. 主持山东省自然科学基金青年基金项目：长链脂肪醇氧化酶FAO3和FAO4b参与拟南芥角质组分α,ω二元羧酸合成的功能解析（ZR2020QC018），2021-2023，15万，已结题；

3. 主持中国博士后科学基金面上项目二等资助：长链脂肪醇氧化酶FAO3参与表皮蜡质降解的功能解析（2018M642961），2018-2019，5万，已结题。

四、 教学科研成果（论文、专著、专利等）

研究论文 (第一作者或通讯作者)

1. Li, S^#., Zhang, X^#., Huang, H^#., ... & Yang, X*., Zhao, H*., & Lü, S*. (2025). Deciphering the core shunt mechanism in Arabidopsis cuticular wax biosynthesis and its role in plant environmental adaptation. Nature Plants, 11, 165–175.

2. Yang, X., Huang, H., Wang, Z., ... & Lü, S., & Ma, C., (2025). Acetyl-CoA carboxylase1 influences CER2 activity to mediate the synthesis of very-long-chain fatty acid past C28. Plant Physiology, 197(2): kiae253.

3. Wang, Z., Liu, Y., Huang, H., Zheng, Z., Lü, S., Yang, X*., & Ma, C*., (2025). Functional identification of two glycerol-3-phosphate acyltransferase5 homologs from Chenopodium quinoa. Plant Science, 350:112313.

4. Huang, H., Wang, Y., Yang, P., Zhao H, ... Lü, S*., & Yang, X*. (2024). The Arabidopsis cytochrome P450 enzyme CYP96A4 is involved in the wound-induced biosynthesis of cuticular wax and cutin monomers. The Plant Journal, 118(5): 1619-1634.

5. Huang H^#, Yang X^#, Zheng M, Chen Z, Yang Z, Wu P, Jenks MA, Wang G, Feng T, Liu L, Yang P, Lü S*, Zhao H* (2023) An ancestral role for 3-KETOACYL-COA SYNTHASE3 as a negative regulator of plant cuticular wax synthesis. The Plant Cell, 35: 2251-2270.

6. Li S^#, Yang X^#, Huang H, Qiao R, Jenks MA, Zhao H*, Lü S* (2022) Arabidopsis ACYL-ACTIVATING ENZYME 9 (AAE9) encoding an isobutyl-CoA synthetase is a key factor connecting branched-chain amino acid catabolism with iso-branched wax biosynthesis. New Phytologist, 233: 2458-2470.

7. Tariq F^#, Zhao S^#, Ahmad N, Wang P, Shao Q, Ma C*, Yang X* (2022) Overexpression of β-Ketoacyl CoA Synthase 2B.1 from Chenopodium quinoa promotes suberin monomers' production and salt tolerance in Arabidopsis thaliana. International Journal of Molecular Sciences, 23(21), 13204.

8. Yang X^#, Cui L^#, Li S, Ma C, Kosma DK, Zhao H*, Lü S* (2022) Fatty alcohol oxidase 3 (FAO3) and FAO4b connect the alcohol- and alkane-forming pathways in Arabidopsis stem wax biosynthesis. Journal of Experimental Botany, 73: 3018-3029.

9. Yang X, Feng T, Li S, Zhao H, Zhao S, Ma C, Jenks MA, Lü S (2020) CER16 inhibits post-transcriptional gene silencing of CER3 to regulate alkane biosynthesis. Plant Physiology, 182: 1211-1221.

10. Yang X^#, Wang Z^#, Feng T, Li J, Huang L, Yang B, Zhao H, Jenks MA, Yang P*, Lü S* (2018) Evolutionarily conserved function of the sacred lotus (Nelumbo nucifera Gaertn.) CER2-LIKE family in very-long-chain fatty acid elongation. Planta, 248: 715-727.

11. Yang X^#, Zhao H^#, Kosma DK, Tomasi P, Dyer JM, Li R, Liu X, Wang Z, Parsons EP, Jenks MA, Lü S (2017) The acyl desaturase CER17 is involved in producing wax unsaturated primary alcohols and cutin monomers. Plant Physiology, 173: 1109-1124.

合作论文

1. Li X, Su T, Wang X, Liu Y, Ge J, Huo P, Zhao Y, Wang T, Yu H, Duan M, Jia Y, Yang X, Wang P, Gong Q, Liu J, Ma C (2025) Blocking constitutive autophagy rescues the loss of acquired heat resistance in Arabidopsis fes1a. New Phytologist, 245(6):2569-2583.

2. Su Y, Feng T, Liu CB, Huang H, Wang YL, Fu X, Han ML, Zhang X, Huang X, Wu JC, Song T, Shen H, Yang X, Xu L, Lü S, Chao DY (2023) The evolutionary innovation of root suberin lamellae contributed to the rise of seed plants. Nature Plants, 9(12):1968-1977.

3. Huang H, Yang X, Zheng M, Lü S, Zhao H (2023) Fine-tuning the activities of β-KETOACYL-COA SYNTHASE 3 (KCS3) and KCS12 in Arabidopsis is essential for maintaining cuticle integrity. Journal of Experimental Botany, 74(21):6575-6587.

4. Wang H, Lu Z, Xu Y, Zhang J, Han L, Chai M, Wang ZY, Yang X, Lü S, Tong J, Xiao L, Wen J, Mysore KS, Zhou C (2023) Roles of very-long-chain fatty acids in compound leaf patterning in Medicago truncatula. Plant Physiology, 191: 1751-1770.

5. Yang Q, Yang X, Wang L, Zheng B, Cai Y, Ogutu CO, Zhao L, Peng Q, Liao L, Zhao Y, Zhou H, Han Y (2022) Two R2R3-MYB genes cooperatively control trichome development and cuticular wax biosynthesis in Prunus persica. New Phytologist, 234: 179-196.

6. Wang Y, Yang X, Chen Z, Zhang J, Si K, Xu R, He Y, Zhu F, Cheng Y (2022) Function and transcriptional regulation of CsKCS20 in the elongation of very-long-chain fatty acids and wax biosynthesis in Citrus sinensis flavedo. Horticulture Research 9, uhab027.

7. Huang H, Ayaz A, Zheng M, Yang X, Zaman W, Zhao H, Lü S (2022) Arabidopsis KCS5 and KCS6 play redundant roles in wax synthesis. International Journal of Molecular Sciences, 23(8), 4450.

8. Zhao Z, Yang X, Lü S, Fan J, Opiyo S, Yang P, Mangold J, Mackey D, Xia Y (2020) Deciphering the novel role of AtMIN7 in cuticle formation and defense against the bacterial pathogen infection. International Journal of Molecular Sciences, 21(15), 5547.

9. Zhang P, Wang R, Yang X, Ju Q, Li W, Lü S, Tran LP, Xu J (2020) The R2R3-MYB transcription factor AtMYB49 modulates salt tolerance in Arabidopsis by modulating the cuticle formation and antioxidant defence. Plant, Cell and Environment, 43: 1925-1943.

10. Tang J, Yang X, Xiao C, Li J, Chen Y, Li R, Li S, Lü S, Hu H (2020) GDSL lipase occluded stomatal pore 1 is required for wax biosynthesis and stomatal cuticular ledge formation. New Phytologist, 228: 1880-1896.

五、 联系方式

山东师范大学（长清湖校区）博物楼308室

邮箱：yangxp2006@sdnu.edu.cn